Excerpt from The Argasidae of North America, Central America and Cuba

The taxonomic study of the Argasidae has been beset with difficult problems, some of which are still unsolved. The principal difficulty has been and still is the lack of adequate generic characters. Some of the earlier species were described from very few specimens and frequently with very little knowl edge Oi their biologies. By 1908, Neumann and Nuttall had notably improved the situation by better generic definitions and the use of new specific characters. Both retained as genera only Argas and Ornithodoros. In 1912 Banks added the genus Otobius, and in 1942 Antricola was added by the authors.

In 1908 only some twenty species were known. Now in North America alone there are at least twenty-five species and about thirty new species have been described from other countries. The characters exhibited by these new species have served to clarify certain of the taxonomic problems; they have made others more confusing. Some specific distinctions have found corrobora tive support in the biologies. With the increasing number of new species, it has become more and more difficult to separate Arga; and Ornithodoros and we have found it necessary to change the previous conceptions of these genera. However, no genera have been synonymized. It is probable that subsequent workers will find Others necessary but it is earnestly hoped that none will be erected until reliable generic criteria have been clearly established. The present names serve the purpose and should be changed only for permanent reasons.

In a previous paper1 the senior author emphasized the importance of variation within species. This occurs principally in the ornamentation, and the morphology is reasonably constant. In the Argasidae it is perhaps too early to evaluate fully the variation within species, but at least it is evident that it is of little importance in the genus Ornithodoros. Ornamentation is absent in the known species of the Argasidae except in Ornithodoros coriaceus.

Where confusion has arisen in specific identities, it has been due largely to overlooking characters in the mammillae and hypostome, some Of which are not very easily detected, and the very distinct ones found in the larval stage which heretofore has not been thoroughly studied.

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